生态环境学报 ›› 2021, Vol. 30 ›› Issue (7): 1353-1359.DOI: 10.16258/j.cnki.1674-5906.2021.07.003
所属专题: 生物多样性专题汇编
赵丽1,2(), 郭春燕3, 张文军1, 王晓江1, 刘平生1,2,*(
)
收稿日期:
2021-03-23
出版日期:
2021-07-18
发布日期:
2021-10-09
通讯作者:
*刘平生(1961年生),男,研究员,研究方向为林木引种繁育与生态学。E-mail: nmlkylps@163.com作者简介:
赵丽(1982年生),女,副研究员,研究方向为林木引种繁育与生态学。E-mail: 591518088@qq.com
基金资助:
ZHAO Li1,2(), GUO Chunyan3, ZHANG Wenjun1, WANG Xiaojiang1, LIU Pingsheng1,2,*(
)
Received:
2021-03-23
Online:
2021-07-18
Published:
2021-10-09
摘要:
为了解天然林的群落特征及其相关性,提升天然林的管理及生态防护功能,以大兴安岭扎兰屯天然落叶松(Larix gmelinii)纯林、白桦(Betula platyphylla)次生林、白桦-落叶松混交林为研究对象,分别对乔木层、灌木层和草本层的高度、胸径、冠幅、盖度等指标进行调查,分析群落结构特征,计算物种丰富度指数、多样性指数和均匀度指数,并进行主成分分析(PCA)。结果显示,(1)3个群落中落叶松林的乔木层树高、胸径和灌木层的地径、盖度均最高,草本层特征差异不显著(P>0.05)。(2)落叶松林、白桦-落叶松林中平榛(Corylus heterophylla)和土庄绣线菊(Spiraea pubescens)的重要值均明显高于其他灌木树种;白桦林灌木层只有兴安柳(Salix hsinganica);3种林分草本层均以兴安苔草(Carex chinganensis)为优势种。(3)乔木层、灌木层的丰富度指数、Simpson指数和Shannon-Wiener指数均表现出:白桦-落叶松林>落叶松林>白桦次生林;草本层以落叶松林最为丰富,其Simpson指数、Shannon-Wiener指数均以白桦林为最大;乔木层、灌木层、草本层的Pielou指数均以白桦-落叶松林为最大。(4)PCA排序结果显示,随着林分郁闭度、乔木树高、胸径及灌木盖度、高度、基径的增大,灌木层植物的多样性指数和均匀度增加明显,草本层多样性降低;草本高度越高、盖度越大,草本层多样性指数、均匀度越高。研究表明:乔木层郁闭度、灌木层生长特征(盖度、高度和基径)是影响物种多样性的主因,林分群落特征与植物多样性存在相关性。
中图分类号:
赵丽, 郭春燕, 张文军, 王晓江, 刘平生. 扎兰屯地区典型天然林群落特征及其相关性分析[J]. 生态环境学报, 2021, 30(7): 1353-1359.
ZHAO Li, GUO Chunyan, ZHANG Wenjun, WANG Xiaojiang, LIU Pingsheng. Community Characteristics and Their Correlation Analysis of Typical Natural Forest in Zhalantun[J]. Ecology and Environment, 2021, 30(7): 1353-1359.
层 Layer | 特征参数 Characteristic parameter | 落叶松林 Larch forest | 白桦-落叶松林 Birch-larch forest | 白桦林 Birch forest |
---|---|---|---|---|
乔木层 Arbor | 树高Tree height/m | 13.87±1.22a | 10.78±0.47b | 6.97±0.20c |
胸径 Tree diameter breast height/cm | 18.73±4.63a | 13.86±0.45a | 7.24±0.37b | |
郁闭度 Tree canopy closure/% | 97.51±4.30a | 87.78±17.80a | 56.92±2.92b | |
灌木层 Shrub | 株高 Shrub height/m | 0.66±0.09a | 0.7±0.04a | 0.46±0.06b |
地径 Shrub ground diameter/cm | 0.67±0.1a | 0.61±0.13a | 0.38±0.16a | |
盖度 Shrub coverage/% | 55.78±31.31a | 39.25±9.5ab | 10.33±4.33b | |
冠幅 Shrub crown width/cm | 31.19±11.74a | 28.92±7.96a | 20.00±10.33a | |
草本层 Herb | 多度 Herb coverage/plant | 12.24±5.92a | 8.44±2.04a | 7.00±1.20a |
高度 Herb height/cm | 22.55±7.69a | 14.50±1.71a | 29.52±16.77a | |
盖度 Herb coverage/% | 60.65±26.55a | 38.52±2.86a | 56.32±2.63a |
表1 3种林分乔木层、灌木层、草本层特征值比较
Table 1 Comparison of characteristic values of tree layer, shrub layer and herb layer in three forest types
层 Layer | 特征参数 Characteristic parameter | 落叶松林 Larch forest | 白桦-落叶松林 Birch-larch forest | 白桦林 Birch forest |
---|---|---|---|---|
乔木层 Arbor | 树高Tree height/m | 13.87±1.22a | 10.78±0.47b | 6.97±0.20c |
胸径 Tree diameter breast height/cm | 18.73±4.63a | 13.86±0.45a | 7.24±0.37b | |
郁闭度 Tree canopy closure/% | 97.51±4.30a | 87.78±17.80a | 56.92±2.92b | |
灌木层 Shrub | 株高 Shrub height/m | 0.66±0.09a | 0.7±0.04a | 0.46±0.06b |
地径 Shrub ground diameter/cm | 0.67±0.1a | 0.61±0.13a | 0.38±0.16a | |
盖度 Shrub coverage/% | 55.78±31.31a | 39.25±9.5ab | 10.33±4.33b | |
冠幅 Shrub crown width/cm | 31.19±11.74a | 28.92±7.96a | 20.00±10.33a | |
草本层 Herb | 多度 Herb coverage/plant | 12.24±5.92a | 8.44±2.04a | 7.00±1.20a |
高度 Herb height/cm | 22.55±7.69a | 14.50±1.71a | 29.52±16.77a | |
盖度 Herb coverage/% | 60.65±26.55a | 38.52±2.86a | 56.32±2.63a |
层 layer | 植物种 Plant species | 重要值 Importance value/% | ||
---|---|---|---|---|
落叶 松林 Larch forest | 白桦-落叶松林 Birch-larch forest | 白桦林 Birch forest | ||
乔木层Tree | 落叶松 Larix gmelinii | 70.40 | 44.74 | 11.69 |
山杨 Populus davidiana | 9.71 | 2.95 | 5.83 | |
白桦 Betula platyphylla | 9.87 | 22.53 | 76.62 | |
柞木 Xylosma racemosum | 5.24 | 17.35 | ‒ | |
黑桦 Betula dahurica | 4.78 | 12.43 | 5.86 | |
灌木层Shrub | 土庄绣线菊 Spiraea pubescens | 28.72 | 35.32 | ‒ |
平榛 Corylus heterophylla | 56.66 | 53.24 | ‒ | |
小叶鼠李 Rhamnus parvifolia | ‒ | 1.95 | ‒ | |
胡枝子 Lespedeza bicolor | 14.62 | 6.98 | ‒ | |
山刺梅 Rosa davurica | ‒ | 2.51 | ‒ | |
兴安柳 Salix hsinganica | ‒ | ‒ | 100.00 | |
草本层Herb | 地榆 Sanguisorba officinalis | 5.44 | 7.46 | 6.58 |
东方草莓 Fragaria orientalis | 16.56 | 10.82 | ‒ | |
兴安苔草 Carex chinganensis | 29.61 | 38.37 | 36.45 | |
蒙古蒿 Mongolian wormwood | 2.22 | ‒ | ‒ | |
唐松草Thalictrum petaloideum | 4.25 | 4.67 | 1.10 | |
鼠掌老鹳草 Geranium sibiricum | 3.31 | 5.07 | 15.41 | |
野火球 Trifolium lupinaster | 2.66 | 2.95 | 0.91 | |
白莲蒿 Artemisia sacrorum | 4.79 | 4.95 | 7.97 | |
龙牙草 Agrimonia pilosa | 1.14 | ‒ | 2.53 | |
黑沙蒿 Artemisia ordosica | 1.17 | 1.56 | ‒ | |
细叶鸢尾 Iris tenuifolia | 4.79 | 3.12 | 3.26 | |
小红菊 Dendranthema chanetii | 7.71 | 4.27 | ‒ | |
茜草 Rubia cordifolia | 1.31 | 1.06 | 4.43 | |
费菜 Sedum aizoon | 3.59 | 1.21 | ‒ | |
冰草 Agropyron cristatum | 5.08 | 3.69 | ‒ | |
东北酸模 Rumex thyrsiflorus | 0.84 | ‒ | 3.73 | |
狗尾草 Setaria viridis | ‒ | 0.86 | 5.37 | |
铃兰 Convallaria majalis | ‒ | 1.05 | ‒ | |
楔叶菊 Dendranthema naktongense | ‒ | 1.19 | ‒ | |
柴胡 Bupleurum chinense | ‒ | 1.17 | ‒ | |
益母草 Leonurus artemisia | ‒ | 1.05 | ‒ | |
橐吾 Ligularia spp. | ‒ | ‒ | 4.17 |
表2 3种林分乔木层、灌木层、草本层重要值比较
Table 2 Comparison of important values for three stands of arbor layer, shrub layer and herb layer
层 layer | 植物种 Plant species | 重要值 Importance value/% | ||
---|---|---|---|---|
落叶 松林 Larch forest | 白桦-落叶松林 Birch-larch forest | 白桦林 Birch forest | ||
乔木层Tree | 落叶松 Larix gmelinii | 70.40 | 44.74 | 11.69 |
山杨 Populus davidiana | 9.71 | 2.95 | 5.83 | |
白桦 Betula platyphylla | 9.87 | 22.53 | 76.62 | |
柞木 Xylosma racemosum | 5.24 | 17.35 | ‒ | |
黑桦 Betula dahurica | 4.78 | 12.43 | 5.86 | |
灌木层Shrub | 土庄绣线菊 Spiraea pubescens | 28.72 | 35.32 | ‒ |
平榛 Corylus heterophylla | 56.66 | 53.24 | ‒ | |
小叶鼠李 Rhamnus parvifolia | ‒ | 1.95 | ‒ | |
胡枝子 Lespedeza bicolor | 14.62 | 6.98 | ‒ | |
山刺梅 Rosa davurica | ‒ | 2.51 | ‒ | |
兴安柳 Salix hsinganica | ‒ | ‒ | 100.00 | |
草本层Herb | 地榆 Sanguisorba officinalis | 5.44 | 7.46 | 6.58 |
东方草莓 Fragaria orientalis | 16.56 | 10.82 | ‒ | |
兴安苔草 Carex chinganensis | 29.61 | 38.37 | 36.45 | |
蒙古蒿 Mongolian wormwood | 2.22 | ‒ | ‒ | |
唐松草Thalictrum petaloideum | 4.25 | 4.67 | 1.10 | |
鼠掌老鹳草 Geranium sibiricum | 3.31 | 5.07 | 15.41 | |
野火球 Trifolium lupinaster | 2.66 | 2.95 | 0.91 | |
白莲蒿 Artemisia sacrorum | 4.79 | 4.95 | 7.97 | |
龙牙草 Agrimonia pilosa | 1.14 | ‒ | 2.53 | |
黑沙蒿 Artemisia ordosica | 1.17 | 1.56 | ‒ | |
细叶鸢尾 Iris tenuifolia | 4.79 | 3.12 | 3.26 | |
小红菊 Dendranthema chanetii | 7.71 | 4.27 | ‒ | |
茜草 Rubia cordifolia | 1.31 | 1.06 | 4.43 | |
费菜 Sedum aizoon | 3.59 | 1.21 | ‒ | |
冰草 Agropyron cristatum | 5.08 | 3.69 | ‒ | |
东北酸模 Rumex thyrsiflorus | 0.84 | ‒ | 3.73 | |
狗尾草 Setaria viridis | ‒ | 0.86 | 5.37 | |
铃兰 Convallaria majalis | ‒ | 1.05 | ‒ | |
楔叶菊 Dendranthema naktongense | ‒ | 1.19 | ‒ | |
柴胡 Bupleurum chinense | ‒ | 1.17 | ‒ | |
益母草 Leonurus artemisia | ‒ | 1.05 | ‒ | |
橐吾 Ligularia spp. | ‒ | ‒ | 4.17 |
层 Layer | 物种多样性指数 Species diversity index | 落叶松林 Larch forest | 白桦-落叶松混交林 Birch-larch forest | 白桦林 Birch forest |
---|---|---|---|---|
乔木层 Arbor | 丰富度R | 2.67±1.53ab | 4.33±0.58a | 2.00±0b |
Simpson指数D | 0.21±0.20b | 0.67±0.01a | 0.13±0.18b | |
Shannon-Wienner 指数H | 0.43±0.40b | 1.25±0.03a | 0.22±0.26b | |
Pielou指数1 Jsw | 0.31±0.29b | 0.86±0.05a | 0.29±0.33b | |
Pielou指数2 Jsi | 0.28±0.26a | 0.88±0.04a | 0.28±0.40a | |
Alatalo指数Ea | 0.33±0.29b | 0.83±0.06a | 0.49±0.21ab | |
灌木层 Shrub | 丰富度R | 3.00±0a | 3.33±1.53a | 1.00±0b |
Simpson指数D | 0.36±0.08a | 0.59±0.34a | 0.00b | |
Shannon-Wienner 指数H | 0.63±0.14a | 0.62±0.20a | 0.00b | |
Pielou指数1 Jsw | 0.58±0.13a | 0.58±0.19a | 0.00b | |
Pielou指数2 Jsi | 0.53±0.13ab | 0.90±0.49a | 0.00b | |
Alatalo指数Ea | 0.63±0.04a | 0.65±0.11a | 0.00b | |
草本层 Herb | 丰富度R | 16.00±2.00a | 15.67±2.08a | 10.67±2.08b |
Simpson指数D | 0.77±0.07a | 0.78±0.07a | 0.84±0.02a | |
Shannon-Wienner 指数H | 1.79±0.24a | 1.96±0.29a | 2.04±0.10a | |
Pielou指数1 Jsw | 0.65±0.07b | 0.71±0.07ab | 0.87±0.03a | |
Pielou指数2 Jsi | 0.82±0.07a | 0.84±0.07a | 0.93±0.01a | |
Alatalo指数Ea | 0.68±0.06ab | 0.62±0.05b | 0.77±0.02a |
表3 3种典型林分多样性指数、均匀度和丰富度指数比较
Table 3 Comparison of diversity index, evenness index and richness index of three typical stands
层 Layer | 物种多样性指数 Species diversity index | 落叶松林 Larch forest | 白桦-落叶松混交林 Birch-larch forest | 白桦林 Birch forest |
---|---|---|---|---|
乔木层 Arbor | 丰富度R | 2.67±1.53ab | 4.33±0.58a | 2.00±0b |
Simpson指数D | 0.21±0.20b | 0.67±0.01a | 0.13±0.18b | |
Shannon-Wienner 指数H | 0.43±0.40b | 1.25±0.03a | 0.22±0.26b | |
Pielou指数1 Jsw | 0.31±0.29b | 0.86±0.05a | 0.29±0.33b | |
Pielou指数2 Jsi | 0.28±0.26a | 0.88±0.04a | 0.28±0.40a | |
Alatalo指数Ea | 0.33±0.29b | 0.83±0.06a | 0.49±0.21ab | |
灌木层 Shrub | 丰富度R | 3.00±0a | 3.33±1.53a | 1.00±0b |
Simpson指数D | 0.36±0.08a | 0.59±0.34a | 0.00b | |
Shannon-Wienner 指数H | 0.63±0.14a | 0.62±0.20a | 0.00b | |
Pielou指数1 Jsw | 0.58±0.13a | 0.58±0.19a | 0.00b | |
Pielou指数2 Jsi | 0.53±0.13ab | 0.90±0.49a | 0.00b | |
Alatalo指数Ea | 0.63±0.04a | 0.65±0.11a | 0.00b | |
草本层 Herb | 丰富度R | 16.00±2.00a | 15.67±2.08a | 10.67±2.08b |
Simpson指数D | 0.77±0.07a | 0.78±0.07a | 0.84±0.02a | |
Shannon-Wienner 指数H | 1.79±0.24a | 1.96±0.29a | 2.04±0.10a | |
Pielou指数1 Jsw | 0.65±0.07b | 0.71±0.07ab | 0.87±0.03a | |
Pielou指数2 Jsi | 0.82±0.07a | 0.84±0.07a | 0.93±0.01a | |
Alatalo指数Ea | 0.68±0.06ab | 0.62±0.05b | 0.77±0.02a |
群落特征参数 Community characteristic parameter | PCA | |||
---|---|---|---|---|
Axis 1 | Axis 2 | Axis 3 | Axis 4 | |
特征值 Eigenvalues | 0.6842 | 0.2037 | 0.0398 | 0.0388 |
累积贡献率 Explained variation (cumulative) | 68.42 | 88.79 | 92.77 | 96.65 |
表4 群落特征PCA排序特征值
Table 4 Community characteristics PCA ordering eigenvalues
群落特征参数 Community characteristic parameter | PCA | |||
---|---|---|---|---|
Axis 1 | Axis 2 | Axis 3 | Axis 4 | |
特征值 Eigenvalues | 0.6842 | 0.2037 | 0.0398 | 0.0388 |
累积贡献率 Explained variation (cumulative) | 68.42 | 88.79 | 92.77 | 96.65 |
图1 林分群落特征对植物多样性影响的二维排序图 箭头表示林分特征The arrow indicats the community characteristics:(1)TH:tree height,乔木树高;TDBH:tree diameter breast height,乔木胸径;TCC:Tree canopy closure,乔木郁闭度;(2)SH:shrub height,灌木株高;SC:hrub coverage,灌木盖度;SGD:shrub ground diameter,灌木地径;(3)HC:herb coverage,草本盖度;HH:herb height,草本高度。 实心质点表示多样性指数The solid particle indicats the diversity index: (1)T-R:Richnessindex of tree,乔木丰富度;T-D:Simpson index of tree,乔木Simpson指数;T-H′:Shannon-wiener index of tree,乔木Shannon-wiener指数;T-Jsw:Pielou index 1 of tree,乔木Pielou指数1;T-Jsi:Pielou index 2 of tree,乔木Pielou指数2;T-Ea:Alatalo index of tree,乔木Alatalo指数; (2)S-R:Richness index of shrub,灌木丰富度;S-D:Simpson index of shrub,灌木Simpson指数;S-H′:Shannon-wiener index of shrub,灌木Shannon-wiener指数;S-Jsw:Pielou index 1 of shrub,灌木Pielou 指数1:S-Jsi:Pielou index 2 of shrub,灌木Pielou指数2;S-Ea:Alatalo index of shrub,灌木Alatalo指数; (3)H-R:Richness index of herb,草本丰富度;H-D:Simpson index of herb,草本Simpson指数;H-H′:Shannon-wiener index of herb,草本Shannon-wiener指数;H-Jsw:Pielou index 1 of herb,草本Pielou指数1;H-Jsi:Pielou index 2 of herb,草本Pielou指数2;H-Ea:Alatalo index ofherb,草本Alatalo指数
Fig. 1 Two dimensional sequence diagram for the effect of stand community characteristics on plant diversity
[1] |
CADOTTE M W, CARSCADDEN K, MIROTCHNICK N, 2011. Beyond species functional diversity and the maintenance of ecological processes and services[J]. Journal of Applied Ecology, 48(5): 1079-1087.
DOI URL |
[2] |
LEVINE J M, HILLERISLAMBERS J, 2009. The importance of niches for the maintenance of species diversity[J]. Nature, 461(7261): 254-257.
DOI URL |
[3] |
LI L, HUAR IG Z L, YE W H, et al., 2009. Spatial distributions of tree species in a subtropical forest of china[J]. Oikos, 118(4): 495-502.
DOI URL |
[4] |
TRENTINI C P, CAMPANELLO P I, VILLAGRA M, et al., 2017. Thinning of loblolly pine plantations in subtropical Argentina: Impact on microclimate and understory vegetation[J]. Forest Ecology and Management, 384: 236-247.
DOI URL |
[5] |
WANG Q G, BAO D C, GUO Y L, et al., 2014. Species associations in a species-rich subtropical forest were not well-explained by stochastic geometry of biodiversity[J]. PLoS One, 9(5): e97300.
DOI URL |
[6] | 崔佳佳, 铁牛, 2021. 大兴安岭北部森林群落结构及植物多样性特征研究[J]. 西北林学院学报, 36(2): 24-30. |
CUI J J, TIE N, 2021. Forest community structureand plant diversity characteristics in Northern Greater Khingan Mountains[J]. Journal of Northwest Forestry University, 36(2): 24-30. | |
[7] | 杜志, 亢新刚, 孟京辉, 等, 2013. 长白山杨桦次生林主要树种的空间分布格局及其关联性[J]. 东北林业大学学报, 41(4): 36-42. |
DU Z, KANG X G, M J H, et al., 2013. Spatial distribution patterns and associations of dominant tree species in poplar-birch secondary forest stand in Changbai Mountains[J]. Journal of Northeast Forestry University, 41(4): 36-42. | |
[8] |
桂旭君, 练琚愉, 张入匀, 等, 2019. 鼎湖山南亚热带常绿阔叶林群落垂直结构及其物种多样性特征[J]. 生物多样性, 27(6): 619-629.
DOI |
GUI X J, LIAN J Y, ZHANG R Y, et al., 2019. Vertical structure and its biodiversity in a subtropical evergreen broad leaved forest at Dinghushan in Guangdong Province, China[J]. Biodiversity Science, 27(6): 619-629.
DOI URL |
|
[9] | 蒋蕾, 刘兆刚, 2014. 大兴安岭天然白桦林群落结构特征及其物种多样性[J]. 森林工程, 30(4): 12-17. |
JIANG L, LIU Z G, 2014. Structure characteristics of natural birch community and species diversity in Great Xing’an mountains[J]. Forest Engineering, 30(4): 12-17. | |
[10] | 雷相东, 2000. 东北过伐林区几种典型森林类型的物种和林分结构多样性及采伐的影响研究[D]. 北京: 北京林业大学. |
LEI X D, 2000. Studies on plant species diversity and stand structural diversity of typical forest types and cutting effects on them in over-logged forests in Northeastern China[D]. Beijing: Beijing Forestry University. | |
[11] | 李冰, 2009. 大兴安岭兴安落叶松林健康评价研究[D]. 北京: 北京林业大学. |
LI B, 2009. Research on forest health evaluation of Larix gmelin in the great Xing’an Mountains[D]. Beijing: Beijing Forestry University. | |
[12] | 黎芳, 潘萍, 宁金魁, 等, 2016. 飞播马尾松林林分空间结构对林下植被多样性的影响[J]. 东北林业大学学报, 44(1): 31-35, 40. |
LI F, PAN P, NING J K, et al., 2016. Effects of stand spatial structure on understory vegetation diversity of aerial seeding Pinus massoniana plantations[J]. Journal of Northeast Forestry University, 44(11): 31-35, 40. | |
[13] | 连灵燕, 张海珠, 2012. 白桦天然次生林种群种间关联及演替规律的研究[J]. 山西师范大学学报(自然科学版), 26(4): 62-65. |
LIAN L Y, ZHANG H Z, 2012. Study on interspecific association and community succession of Betula platypylla natural secondary forest[J]. Shanxi Normal University (Science), 26(4): 62-65. | |
[14] | 刘林馨, 2012. 小兴安岭森林生态系统植物多样性及生态服务功能价值研究[D]. 哈尔滨:东北林业大学. |
LIU L X, 2012. Study on the palnt biodiversity and ecosystem service functions evaluation of forest ecosystem in Xiaoxing’an mountains[D]. Harbin: Northeast Forest University. | |
[15] | 刘召强, 刘艺平, 冯理明, 等, 2020. 额尔齐斯河流域垂枝桦林群落结构特征与物种多样性相关分析[J]. 西北植物学报, 40(3): 532-542. |
LIU Z Q, LIU Y P, FENG L M, et al., 2020. Differences in community characteristics, species diversity, and their coupling associations among Betula pendula forests in Irtysh river watershed[J]. Acta Botanica Boreali-Occidentalia Sinica, 40(3): 532-542. | |
[16] |
马文静, 张庆, 牛建明, 等, 2013. 物种多样性和功能群多样性与生态系统生产力的关系——以内蒙古短花针茅草原为例[J]. 植物生态学报, 37(7): 620-630.
DOI |
MA W J, ZHANG Q, NIU J M, et al., 2013. Relationship of ecosystem primary productivity to species diversity and functional group diversity: Evidence from Stipa breviflora grassland in Nei Mongol[J]. Chinese Journal of Plant Ecology, 37(7): 620-630.
DOI URL |
|
[17] | 曲红, 王百田, 王棣, 等, 2010. 黄土区不同配置人工林物种多样性研究[J]. 生态环境学报, 19(4): 843-848. |
QU H, WANG B T, WANG D, et al., 2010. Research of artificial forest plant diversity under different configuration of loess area[J]. Ecology and Environmental Sciences, 19(4): 843-848. | |
[18] | 矢佳昱, 韩海荣, 程小琴, 等, 2018. 辽河源自然保护区不同年龄天然油松林林下草本层植物物种多样性及其环境解释[J]. 生态学杂志, 37(5): 1326-1333. |
SHI J Y, HAN H R, CHENG X Q, et al., 2018. Environmental factors affecting plant species diversity of understory herbaceous communities in a chronosequence of Pinus tabuliformis forest in Liaoheyuan Nature Reserve[J]. Chinese Journal of Ecology, 37(5): 1326-1333. | |
[19] | 覃志杰, 董威, 刘泰瑞, 等, 2019. 油松天然次生林林下植物多样性对林分密度的响应研究[J]. 山西农业大学学报(自然科学版), 39(1): 61-67. |
TAN Z J, DONG W, LIU T R, et al., 2019. Responses of understory plant diversity to stand density in natural secondary forests of Pinus tabulaeformis [J]. Journal of Shanxi Agricultural University (Natural Science Edition), 39(1): 61-67. | |
[20] | 王玲, 2020. 林分密度对油松人工林群落结构和植物多样性的影响[J]. 生态环境学报, 29(12): 2328-2336. |
WANG L, 2020. Effects of different stand densities on community structure and species diversity of Pinus tabulaeformis plantation[J]. Ecology and Environmental Sciences, 29(12): 2328-2336. | |
[21] | 邢晖, 2014. 大兴安岭落叶松白桦混交林林分空间结构优化技术研究[D]. 哈尔滨: 东北林业大学. |
XING H, 2014. Study on the forest stand spatial structure optimization technology for Larch-Betula platyphylla mixed forest stand in Daxing’an mountains[D]. Harbin: Northeast Forest University. | |
[22] | 赵淑清, 方精云, 朴世龙, 等, 2004. 大兴安岭呼中地区白卡鲁山植物群落结构及其多样性研究[J]. 生物多样性, 12(1): 182-189. |
ZHAO S Q, FANG J Y, PIAO S L, et al., 2004. Structure and species diversity of boreal forests in Mt. Baikalu, Huzhong area, Daxing’an Mountains, Northeast China[J]. Biodiversity Science, 12(1): 182-189. | |
[23] | 张建宇, 2018. 大兴安岭森林植物多样性、群落结构特征及耦合性关系分析[D]. 哈尔滨: 东北林业大学. |
ZHANG J Y, 2018. Characteristics of plant diversity and community structure in Daxing’anling forests and association analysis[D]. Harbin: Northeast Forest University. | |
[24] | 张佳, 李生宇, 靳正忠, 等, 2011. 防护林下草本植物层片物种多样性与环境因子的关系[J]. 干旱区研究, 28(1): 118-125. |
ZHANG J, LI S Y, JIN Z Z, et al., 2011. Relationship between species diversity of herbaceous plants in the shelterbelt and environment factors[J]. Arid Zone Research, 28(1): 118-125.
DOI URL |
|
[25] | 张建宇, 王文杰, 杜红居, 等, 2018. 大兴安岭呼中地区3种林分的群落特征、物种多样性差异及其耦合关系[J]. 生态学报, 38(13): 4684-4693. |
ZHANG J Y, WANG W J, DUAN H J, et al., 2018. Differences in community characteristics,species diversity and their coupling associations among three forest types in the Huzhongarea Daxinganling Mountains[J]. Acta Ecologica Sinica, 38(13): 4684-4693. | |
[26] | 张维伟, 薛文艳, 杨斌, 等, 2019. 桥山栎林群落结构特征与物种多样性相关关系分析[J]. 生态学报, 39(11): 3991-4001. |
ZHANG W W, XUE W Y, YANG B, et al., 2019. Differences incommunity characteristics,species diversity, and their coupling associations among three Quercus acutissima forests in Qiaoshan[J]. Acta Ecologica Sinica, 39(11): 3991-4001. | |
[27] | 周晓果, 温远光, 朱宏光, 等, 2017. 大明山常绿阔叶林冠层垂直结构与林下植物更新[J]. 应用生态学报, 28(2): 367-374. |
ZHOU X G, WEN Y G, ZHU H G, et al., 2017. Canopy vertical structure and understory plant regeneration of an evergreen broadleaved forest in Damingshan, Guangxi, China[J]. Chinese Journal of Applied Ecology, 28(2): 367-374. | |
[28] | 张柳桦, 齐锦秋, 李婷婷, 等, 2019. 林分密度对新津文峰山马尾松人工林林下物种多样性和生物量的影响[J]. 生态学报, 39(15): 5709-5717. |
ZHANG L H, QI J Q, LI T T, et al., 2019. Effects of stand density on understory plant diversity and biomass in a Pinus massoniana plantation in Wenfeng Mountain, Xinjin County[J]. Acta Ecologica Sinica, 39(15): 5709-5717. |
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