南亚热带常绿阔叶林植物不同分类水平上的空间分布格局及空间关联

doi:10.16258/j.cnki.1674-5906.2022.03.002

生态环境学报 ›› 2022, Vol. 31 ›› Issue (3): 440-450.DOI: 10.16258/j.cnki.1674-5906.2022.03.002

南亚热带常绿阔叶林植物不同分类水平上的空间分布格局及空间关联

温智峰¹(), 魏识广², 李林¹^,²^,^*(), 叶万辉³^,⁴, 练琚愉³^,⁴

1.桂林电子科技大学生命与环境科学学院,广西桂林 541004
2.广西师范大学珍稀濒危动植物生态与环境保护教育部重点实验室,广西桂林 541006
3.中国科学院华南植物园退化生态系统植被恢复与管理重点实验室,广东广州 510650
4.中国科学院核心植物园植物生态中心,广东广州 510650

收稿日期:2021-11-10 出版日期:2022-03-18 发布日期:2022-05-25
通讯作者: *李林（1978年生）,女,教授,博士,从事生物多样性分布格局与保护研究。E-mail: dhslilin@scbg.ac.cn
作者简介:温智峰（1997年生）,男,硕士研究生,从事生物多样性与环境生态学研究。E-mail: wen1059540044@163.com
基金资助:
国家自然科学基金项目(32060305);广西自然科学基金面上基金项目(2020GXNSFAA159108);广西师范大学珍稀濒危动植物生态与环境保护教育部重点实验室研究基金项目(ERESEP2021Z06)

Spatial Distribution Patterns and Spatial Associations of Evergreen Broad-leaved Forest Plants in Tropical South Asia at Different Taxonomic Levels

WEN Zhifeng¹(), WEI Shiguang², LI Lin¹^,²^,^*(), YE Wanhui³^,⁴, LIAN Juyu³^,⁴

1. College of Life and environmental Science, Guilin University of Electronic Technology, Guilin 541004, P. R. China
2. College of Life Science, Guangxi Normal University, Key Laboratory of Ecology of Rare and Endangered Species and Environmental Protection, Ministry of Education, Guilin 541006, P. R. China
3. Key Laboratory of Vegetation Restoration and Management of South China Botanical Garden, Chinese Academy of Sciences, Guangzhou 510650, P. R. China
4. Center of Plant Ecology, Core Botanical Gardens, Chinese Academy of Sciences, Guangzhou 510650, P. R. China

Received:2021-11-10 Online:2022-03-18 Published:2022-05-25

摘要/Abstract

摘要：

为探讨中国南亚热带常绿阔叶林植物在科、属、种不同分类水平上空间分布格局、空间关联的共性规律及其成因,加深对中国南亚热带区域群落生态学特性的了解。选取中国南亚热带区域4个不同纬度样地中个体数量排名前3的优势科为研究对象（DBH,diameter at breast height≥1 cm）,采用L(t) 方程统计科、属、种空间分布格局,并以空间聚集程度指标L₁₀比较各样地科、属、种水平的聚集程度,采用L₁₂(t) 方程分析各样地科、属、种间个体的空间关联。结果表明,（1）聚集分布是南亚热带常绿阔叶林群落植物在科、属、种3个分类水平上的最主要空间分布格局类型;各样地在不同分类水平上的聚集分布比例随纬度梯度变化未表现出明显规律,但均表现出分布格局的尺度依赖性,即随着空间尺度增大,聚集分布比例下降,随机分布比例上升这一共性规律。（2）各样地空间聚集程度L₁₀均表现出随科→属→种分类水平细化而增大的规律。（3）独立不相关为各样地不同分类水平空间关联的最主要类型;独立不相关所占比例在科、属、种3个水平的变化规律不一致,但在物种水平,各样地3类亲缘远近的独立不相关占各类种间关联比例表现出同科同属占比>不同科不同属占比>同科不同属占比这一共性规律。（4）亲缘个体在种、属水平表现出独立不相关占比随纬度从高到低逐渐减小的趋势,而在科水平统计数据未能体现此规律。综上,植物分类水平的细化一定程度上可以加强空间聚集程度,亲缘关系越近的个体越趋向于聚集分布;不同分类水平上亲缘个体的空间分布格局及关联性,可以真实地反映不同亲缘关系远近群体对生境的适应性及资源利用特点,亲缘关系是影响群落中物种共存和分布的重要因素。

关键词: 空间分布格局, 空间关联, 点格局分析, 南亚热带森林群落, 不同分类水平

Abstract:

This study investigated the common patterns and causes of spatial distribution and association of evergreen broad-leaved forest plants at different taxonomic levels from families to species in the south subtropics of China and expands our understanding of the ecological characteristics of regional communities in the south subtropics of China. In this study, the top three dominant families (DBH, Diameter at breast height ≥1 cm) in the number of individuals in four different latitudinal sample sites in the south subtropical region of China were selected. We used the L(t) function to count the spatial distribution patterns of families, genera and species, and compared the degree of aggregation at the level of families, genera and species in each plot with the spatial aggregation index L₁₀, and used the L₁₂(t) function to analyze the spatial association of individuals among families, genera, and species in each plot. The results showed that (1) aggregation distribution was the most important spatial distribution pattern of plants in the southern subtropical broadleaf evergreen forest communities at three taxonomic levels. The proportion of aggregation distribution at each taxonomic level did not show a clear pattern with latitudinal gradient, but all showed a scale-dependent distribution pattern (i.e., the proportion of aggregation distribution decreased and the proportion of random distribution increased as the spatial scale increased.) (2) The spatial aggregation degree L₁₀ of each sample site increased with the refinement of family→genera→species taxonomic levels. (3) mutual independence was the most dominant type of spatial association at different taxonomic levels at each site. Although the proportion of this type of association varied among the three levels of family, genus and species, it followed the same order (i.e., same family to same genus>different family to different genus>same family to different genus) at the species level at each site. (4) The proportion of independently unrelated individuals at the species and genus levels tended to decrease from higher to lower latitudes, while the statistics at the family level failed to reflect this pattern. In summary, the refinement of plant taxonomic levels can strengthen the degree of spatial aggregation to a certain extent, and individuals with closer affinities tend to be more aggregated and distributed; the spatial distribution patterns and correlations of related individuals at different taxonomic levels can truly reflect the habitat adaptation and resource utilization characteristics of groups with different affinities, and affinities are important factors that influence the coexistence and distribution of species in communities.

Key words: spatial distribution pattern, spatial association, point pattern analysis, southern subtropical forest communities, different taxonomic levels

中图分类号:

Q948
X173

温智峰, 魏识广, 李林, 叶万辉, 练琚愉. 南亚热带常绿阔叶林植物不同分类水平上的空间分布格局及空间关联[J]. 生态环境学报, 2022, 31(3): 440-450.

WEN Zhifeng, WEI Shiguang, LI Lin, YE Wanhui, LIAN Juyu. Spatial Distribution Patterns and Spatial Associations of Evergreen Broad-leaved Forest Plants in Tropical South Asia at Different Taxonomic Levels[J]. Ecology and Environment, 2022, 31(3): 440-450.

图/表 9

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样地 Plot	经度 Longitude	纬度 Latitude	海拔 Altitude/m	科数 Number of families	属数 Number of genera	种数 Number of species	胸径≥1 cm个体总数 Total number of individuals with DBH≥1 cm	多度≥20株物种数 Number of species with abundance≥20
井冈山 Jinggang Mountain	114°11′E	26°23′N	450-500	33	53	108	3947	41
猫儿山 Maoer Mountain	110°30′E	25°56′N	1450-1500	36	65	125	2465	32
南岭 Nanling Mountain	112°75′E	24°39′N	1000-1100	43	83	152	2893	25
鼎湖山 Dinghu Mountain	112°31′E	23°10′N	250-300	38	72	94	3608	24

样地 Plot	经度 Longitude	纬度 Latitude	海拔 Altitude/m	科数 Number of families	属数 Number of genera	种数 Number of species	胸径≥1 cm个体总数 Total number of individuals with DBH≥1 cm	多度≥20株物种数 Number of species with abundance≥20
井冈山 Jinggang Mountain	114°11′E	26°23′N	450-500	33	53	108	3947	41
猫儿山 Maoer Mountain	110°30′E	25°56′N	1450-1500	36	65	125	2465	32
南岭 Nanling Mountain	112°75′E	24°39′N	1000-1100	43	83	152	2893	25
鼎湖山 Dinghu Mountain	112°31′E	23°10′N	250-300	38	72	94	3608	24

样地 Plot	科 Families	属 Genera	种 Species	平均胸径 Average DBH/cm	多度Abundance
井冈山 Jinggang Mountain	杜鹃花科 Ericaceae	杜鹃属 Rhododendron	马银花 Rhododendron ovatum (Lindl.) Planch.	5.03	347
			满山红 Rhododendron mariesii Hemsl. et Wils.	5.09	95
			鹿角杜鹃 Rhododendron latoucheae Franch.	5.95	86
	樟科 Lauraceae	新木姜子属 Neolitsea	浙江新木姜子 Neolitsea var. aurata (Nakai) Yang et P. H. Huang	4.34	38
		润楠属 Machilus	红楠 Machilus thunbergii Sieb. et Zucc.	4.00	64
		润楠属 Machilus	绒毛润楠 Machilus velutina Champ. ex Benth.	2.90	146
		木姜子属 Litsea	石木姜子 Litsea elongata var. faberi (Hemsl.) Yang et P. H. Huang	3.83	356
	山茶科 Theaceae	杨桐属 Adinandra	黄瑞木 Adinandra millettii (Hook. et Arn.) Benth. et Hook. f. ex Hance	10.90	104
		柃木属 Eurya	格药柃 Eurya muricata Dunn	3.07	126
		厚皮香属 Ternstroemia	厚皮香 Ternstroemia gymnanthera (Wight et Arn.) Beddome	6.67	106
		厚皮香属 Ternstroemia	尖萼厚皮香 Ternstroemia luteoflora L. K. Ling	3.29	440
		山茶属 Camellia	长尾毛蕊茶 Camellia caudata Wall.	2.52	21
			短柱茶 Camellia brevistyla (Hayata) Coh. St	2.45	31
			细叶连蕊茶 Camellia euryoides Lindl.	4.75	41
猫儿山 Maoer Mountain	杜鹃花科 Ericaceae	杜鹃属 Rhododendron	厚叶杜鹃 Rhododendron pachyphyllum Fang	5.99	23
			猫儿山杜鹃 Rhododendron maoerense Fang et G. Z. Li	6.45	36
			弯蒴杜鹃 Rhododendron henryi Hance	5.45	51
			映山红 Rhododendron simsii Planch.	3.74	102
	冬青科 Aquifoliaceae	冬青属 Ilex	榕叶冬青 Ilex ficoidea Hemsl.	6.55	221
	冬青科 Aquifoliaceae	冬青属 Ilex	拟榕叶冬青 Ilex subficoides S. Y. Hu	7.82	86
	壳斗科 Fagaceae	栎属 Quercus	麻栎 Quercus acutissima Carr.	8.90	24
		水青冈属 Fagus	水青冈 Fagus longipetiolata Seem.	14.27	328
		锥属 Castanopsis	鹿角锥 Castanopsis lamontii Hance	19.41	56
		青冈属 Cyclobalanopsis	青冈 Cyclobalanopsis glauca (Thunberg) Oersted	10.86	30
			褐叶青冈 Cyclobalanopsis stewardiana (A. Camus) Y. C. Hsu et H. W. Jen	13.33	45
			小叶青冈 Cyclobalanopsis myrsinifolia (Blume) Oersted	10.65	145
南岭 Nanling Mountain	山龙眼科 Proteaceae	山龙眼属 Helicia	网脉山龙眼 Helicia reticulata W. T. Wang	6.96	208
	壳斗科 Fagaceae	柯属 Lithocarpus	短尾柯 Lithocarpus brevicaudatus (Skan) Hayata	6.67	34
		锥属 Castanopsis	黧蒴锥 Castanopsis fissa (Champion ex Bentham) Rehder et E. H. Wilson	9.15	101
		锥属 Castanopsis	米槠 Castanopsis carlesii (Hemsl.) Hayata.	10.84	45
	樟科 Lauraceae	厚壳桂属 Cryptocarya	硬壳桂 Cryptocarya chingii Cheng	6.89	645
		木姜子属 Litsea	广东木姜子 Litsea kwangtungensis H. T. Chang	6.35	54
		山胡椒属 Lindera	广东山胡椒 Lindera kwangtungensis (Liou) Allen	6.04	34
		樟属 Cinnamomum	沉水樟 Cinnamomum micranthum (Hay.) Hay.	7.18	23
		润楠属 Machilus	刨花润楠 Machilus pauhoi Kanehira	9.69	64
			广东润楠 Machilus kwangtungensis Yang	6.70	74
			华润楠 Machilus chinensis (Champ. ex Benth.) Hemsl.	9.17	151
鼎湖山 Dinghu Mountain	大戟科 Euphorbiaceae	野桐属 Mallotus	—	—	—
		血桐属 Macaranga	鼎湖血桐 Macaranga sampsonii Hance	2.85	333
		银柴属 Aporusa	云南银柴 Aporusa yunnanensis (Pax & K. Hoffmann) F. P. Metcalf	5.15	879
	茜草科 Rubiaceae	九节属 Psychotria	九节 Psychotria asiatica Wall.	2.58	165
		茜木属 Aidia	香楠 Aidia canthioides (Champ. ex Benth.) Masam.	2.18	340
		鱼骨木属 Psydrax	鱼骨木 Psydrax dicocca Gaertner	5.28	24
	野牡丹科 Melastomataceae	柏拉木属 Blastus	柏拉木 Blastus cochinchinensis Lour.	1.74	346

样地 Plot	科 Families	属 Genera	种 Species	平均胸径 Average DBH/cm	多度Abundance
井冈山 Jinggang Mountain	杜鹃花科 Ericaceae	杜鹃属 Rhododendron	马银花 Rhododendron ovatum (Lindl.) Planch.	5.03	347
			满山红 Rhododendron mariesii Hemsl. et Wils.	5.09	95
			鹿角杜鹃 Rhododendron latoucheae Franch.	5.95	86
	樟科 Lauraceae	新木姜子属 Neolitsea	浙江新木姜子 Neolitsea var. aurata (Nakai) Yang et P. H. Huang	4.34	38
		润楠属 Machilus	红楠 Machilus thunbergii Sieb. et Zucc.	4.00	64
		润楠属 Machilus	绒毛润楠 Machilus velutina Champ. ex Benth.	2.90	146
		木姜子属 Litsea	石木姜子 Litsea elongata var. faberi (Hemsl.) Yang et P. H. Huang	3.83	356
	山茶科 Theaceae	杨桐属 Adinandra	黄瑞木 Adinandra millettii (Hook. et Arn.) Benth. et Hook. f. ex Hance	10.90	104
		柃木属 Eurya	格药柃 Eurya muricata Dunn	3.07	126
		厚皮香属 Ternstroemia	厚皮香 Ternstroemia gymnanthera (Wight et Arn.) Beddome	6.67	106
		厚皮香属 Ternstroemia	尖萼厚皮香 Ternstroemia luteoflora L. K. Ling	3.29	440
		山茶属 Camellia	长尾毛蕊茶 Camellia caudata Wall.	2.52	21
			短柱茶 Camellia brevistyla (Hayata) Coh. St	2.45	31
			细叶连蕊茶 Camellia euryoides Lindl.	4.75	41
猫儿山 Maoer Mountain	杜鹃花科 Ericaceae	杜鹃属 Rhododendron	厚叶杜鹃 Rhododendron pachyphyllum Fang	5.99	23
			猫儿山杜鹃 Rhododendron maoerense Fang et G. Z. Li	6.45	36
			弯蒴杜鹃 Rhododendron henryi Hance	5.45	51
			映山红 Rhododendron simsii Planch.	3.74	102
	冬青科 Aquifoliaceae	冬青属 Ilex	榕叶冬青 Ilex ficoidea Hemsl.	6.55	221
	冬青科 Aquifoliaceae	冬青属 Ilex	拟榕叶冬青 Ilex subficoides S. Y. Hu	7.82	86
	壳斗科 Fagaceae	栎属 Quercus	麻栎 Quercus acutissima Carr.	8.90	24
		水青冈属 Fagus	水青冈 Fagus longipetiolata Seem.	14.27	328
		锥属 Castanopsis	鹿角锥 Castanopsis lamontii Hance	19.41	56
		青冈属 Cyclobalanopsis	青冈 Cyclobalanopsis glauca (Thunberg) Oersted	10.86	30
			褐叶青冈 Cyclobalanopsis stewardiana (A. Camus) Y. C. Hsu et H. W. Jen	13.33	45
			小叶青冈 Cyclobalanopsis myrsinifolia (Blume) Oersted	10.65	145
南岭 Nanling Mountain	山龙眼科 Proteaceae	山龙眼属 Helicia	网脉山龙眼 Helicia reticulata W. T. Wang	6.96	208
	壳斗科 Fagaceae	柯属 Lithocarpus	短尾柯 Lithocarpus brevicaudatus (Skan) Hayata	6.67	34
		锥属 Castanopsis	黧蒴锥 Castanopsis fissa (Champion ex Bentham) Rehder et E. H. Wilson	9.15	101
		锥属 Castanopsis	米槠 Castanopsis carlesii (Hemsl.) Hayata.	10.84	45
	樟科 Lauraceae	厚壳桂属 Cryptocarya	硬壳桂 Cryptocarya chingii Cheng	6.89	645
		木姜子属 Litsea	广东木姜子 Litsea kwangtungensis H. T. Chang	6.35	54
		山胡椒属 Lindera	广东山胡椒 Lindera kwangtungensis (Liou) Allen	6.04	34
		樟属 Cinnamomum	沉水樟 Cinnamomum micranthum (Hay.) Hay.	7.18	23
		润楠属 Machilus	刨花润楠 Machilus pauhoi Kanehira	9.69	64
			广东润楠 Machilus kwangtungensis Yang	6.70	74
			华润楠 Machilus chinensis (Champ. ex Benth.) Hemsl.	9.17	151
鼎湖山 Dinghu Mountain	大戟科 Euphorbiaceae	野桐属 Mallotus	—	—	—
		血桐属 Macaranga	鼎湖血桐 Macaranga sampsonii Hance	2.85	333
		银柴属 Aporusa	云南银柴 Aporusa yunnanensis (Pax & K. Hoffmann) F. P. Metcalf	5.15	879
	茜草科 Rubiaceae	九节属 Psychotria	九节 Psychotria asiatica Wall.	2.58	165
		茜木属 Aidia	香楠 Aidia canthioides (Champ. ex Benth.) Masam.	2.18	340
		鱼骨木属 Psydrax	鱼骨木 Psydrax dicocca Gaertner	5.28	24
	野牡丹科 Melastomataceae	柏拉木属 Blastus	柏拉木 Blastus cochinchinensis Lour.	1.74	346

样地 Plot	分类水平 Classification levels	尺度 Scale/m
样地 Plot	分类水平 Classification levels	5	10	15	20	25	30	35	40	45	50
井冈山 Jinggang Mountain	科	100	100	100	100	66.7	66.7	66.7	33.3	33.3	33.3
	属	100	100	87.5	87.5	87.5	75.0	75.0	50.0	37.5	37.5
	种	92.9	92.9	85.7	85.7	85.7	85.7	85.7	64.3	57.1	50.0
猫儿山 Maoer Mountain	科	100	100	100	100	66.7	66.7	66.7	33.3	33.3	33.3
	属	66.7	100	100	100	100	100	66.7	50.0	50.0	16.7
	种	83.3	100	100	83.3	83.3	75.0	66.7	50.0	33.3	16.7
南岭 Nanling Mountain	科	100	100	100	100	100	100	100	66.7	33.3	33.3
	属	75.0	100	75.0	87.5	87.5	50.0	37.5	37.5	25.0	12.5
	种	54.5	72.7	63.6	63.6	63.6	45.4	45.4	36.4	18.2	9.1
鼎湖山 Dinghu Mountain	科	100	100	100	100	100	100	100	100	100	100
	属	85.7	100	100	100	100	85.7	85.7	85.7	85.7	85.7
	种	83.3	100	83.3	83.3	100	100	100	100	100	100

南亚热带常绿阔叶林植物不同分类水平上的空间分布格局及空间关联

Spatial Distribution Patterns and Spatial Associations of Evergreen Broad-leaved Forest Plants in Tropical South Asia at Different Taxonomic Levels

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样地 Plot	科及对应L₁₀ Families and its L₁₀	属及对应L₁₀ Genera and its L₁₀	种及对应L₁₀ Species and its L₁₀
井冈山 Jinggang Mountain	杜鹃花科 Ericaceae 2.26	杜鹃属 Rhododendron 2.40	马银花 Rhododendron ovatum (Lindl.) Planch. 2.02
			满山红 Rhododendron mariesii Hemsl. et Wils. 10.75
			鹿角杜鹃 Rhododendron latoucheae Franch. 4.51
	樟科 Lauraceae 1.04	新木姜子属 Neolitsea 6.45	浙江新木姜子 Neolitsea var. aurata (Nakai) Yang et P. H. Huang 11.37
		润楠属 Machilus 3.07	红楠 Machilus thunbergii Sieb. et Zucc. 5.04
		润楠属 Machilus 3.07	绒毛润楠 Machilus velutina Champ. ex Benth. 3.18
		木姜子属 Litsea 1.63	石木姜子 Litsea elongata var. faberi (Hemsl.) Yang et P. H. Huang 1.63
	山茶科 Theaceae 0.72	杨桐属 Adinandra 2.92	黄瑞木 Adinandra millettii (Hook. et Arn.) Benth. et Hook. f. ex Hance 2.92
		柃木属 Eurya 3.66	格药柃 Eurya muricata Dunn 3.51
		厚皮香属 Ternstroemia 1.69	厚皮香 Ternstroemia gymnanthera (Wight et Arn.) Beddome 2.48
		厚皮香属 Ternstroemia 1.69	尖萼厚皮香 Ternstroemia luteoflora L. K. Ling 2.10
		山茶属 Camellia 1.41	长尾毛蕊茶 Camellia caudata Wall. 9.66
			短柱茶 Camellia brevistyla (Hayata) Coh. St 3.92
			细叶连蕊茶 Camellia euryoides Lindl. 5.00
猫儿山 Maoer Mountain	杜鹃花科 Ericaceae 7.54	杜鹃属 Rhododendron 7.77	厚叶杜鹃 Rhododendron pachyphyllum Fang 30.91
			猫儿山杜鹃 Rhododendron maoerense Fang et G. Z. Li 16.25
			弯蒴杜鹃 Rhododendron henryi Hance 26.38
			映山红 Rhododendron simsii Planch. 14.23
	冬青科 Aquifoliaceae 1.85	冬青属 Ilex 1.85	榕叶冬青 Ilex ficoidea Hemsl. 2.06
	冬青科 Aquifoliaceae 1.85	冬青属 Ilex 1.85	拟榕叶冬青 Ilex subficoides S. Y. Hu 3.93
	壳斗科Fagaceae 2.68	栎属 Quercus 7.00	麻栎 Quercus acutissima Carr. 7.00
		水青冈属 Fagus 3.03	水青冈 Fagus longipetiolata Seem. 3.03
		锥属 Castanopsis 3.25	鹿角锥 Castanopsis lamontii Hance 3.18
		青冈属 Cyclobalanopsis 3.50	青冈 Cyclobalanopsis glauca (Thunberg) Oersted 8.33
			褐叶青冈 Cyclobalanopsis stewardiana (A. Camus) Y. C. Hsu et H. W. Jen 4.69
			小叶青冈 Cyclobalanopsis myrsinifolia (Blume) Oersted 6.90
南岭 Nanling Mountain	山龙眼科Proteaceae 5.75	山龙眼属 Helicia 5.75	网脉山龙眼 Helicia reticulata W. T. Wang 6.65
	壳斗科 Fagaceae 7.74	柯属 Lithocarpus 6.61	短尾柯 Lithocarpus brevicaudatus (Skan) Hayata 6.61
		锥属 Castanopsis 9.17	黧蒴锥 Castanopsis fissa (Champion ex Bentham) Rehder et E. H. Wilson 18.92
		锥属 Castanopsis 9.17	米槠 Castanopsis carlesii (Hemsl.) Hayata. 7.04
	樟科 Lauraceae 0.82	厚壳桂属 Cryptocarya 2.12	硬壳桂 Cryptocarya chingii Cheng 2.26
		木姜子属 Litsea 3.94	广东木姜子 Litsea kwangtungensis H. T. Chang 4.18
		山胡椒属 Lindera 3.57	广东山胡椒 Lindera kwangtungensis (Liou) Allen 5.42
		樟属 Cinnamomum 5.98	沉水樟 Cinnamomum micranthum (Hay.) Hay. 6.62
		润楠属 Machilus 1.11	刨花润楠 Machilus pauhoi Kanehira 1.57
			广东润楠 Machilus kwangtungensis Yang 0.93
			华润楠 Machilus chinensis (Champ. ex Benth.) Hemsl. 1.85
鼎湖山 Dinghu Mountain	大戟科 Euphorbiaceae 1.57	野桐属 Mallotus 14.30	—
		血桐属 Macaranga 11.67	鼎湖血桐 Macaranga sampsonii Hance 12.42
		银柴属 Aporusa 1.31	云南银柴 Aporusa yunnanensis (Pax & K. Hoffmann) F. P. Metcalf 1.31
	茜草科 Rubiaceae 4.51	九节属 Psychotria 4.62	九节 Psychotria asiatica Wall. 4.62
		茜木属 Aidia 5.21	香楠 Aidia canthioides (Champ. ex Benth.) Masam. 5.21
		鱼骨木属 Psydrax 6.71	鱼骨木 Psydrax dicocca Gaertner 5.32
	野牡丹科 Melastomataceae 3.24	柏拉木属 Blastus 3.46	柏拉木 Blastus cochinchinensis Lour. 3.46

样地 Plot	同科同属占比 Proportion of the same family and genus/%	不同科不同属占比 Proportion of different families and genera/%	同科不同属占比 Proportion of different genera in the same family/%	占3类种间关联总比例 Proportion in the total of three types of interspecific associations/%
井冈山 Jinggang Mountain	100	96.7	90.9	95.6
猫儿山 Maoer Mountain	100	93.2	75.0	90.9
南岭 Nanling Mountain	100	87.1	85.0	87.3
鼎湖山 Dinghu Mountain	100	100	50.0	73.3

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